Organs

Organs

Fish reproductive organs include testes and ovaries. In most fish species, gonads are paired

organs of similar size, which can be partially or totally fused.[23] There may also be a range of secondary reproductive organs that help in increasing a fish's fitness.
In terms of spermatogonia distribution, the structure of teleosts testes has two types: in the most common, spermatogonia occur all along the seminiferous tubules, while in Atherinomorph fishes they are confined to the distal portion of these structures. Fishes can present cystic or semi-cystic spermatogenesis in relation to the phase of release of germ cells in cysts to the seminiferous tubules lumen.[23]
Fish ovaries may be of three types: gymnovarian, secondary gymnovarian or cystovarian. In the first type, the oocytes are released directly into the coelomic cavity and then enter the ostium, then through the oviduct and are eliminated. Secondary gymnovarian ovaries shed ova into the coelom and then they go directly into the oviduct. In the third type, the oocytes are conveyed to the exterior through the oviduct.[24] Gymnovaries are the primitive condition found in lungfishes, sturgeons, and bowfins. Cystovaries are the condition that characterizes most of the teleosts, where the ovary lumen has continuity with the oviduct.[23] Secondary gymnovaries are found in salmonids and a few other teleosts.
Oogonia development in teleosts fish varies according to the group, and the determination of oogenesis dynamics allows the understanding of maturation and fertilization processes. Changes in the nucleus, ooplasm, and the surrounding layers characterize the oocyte maturation process.[23]
Postovulatory follicles are structures formed after oocyte release; they do not have endocrine function, present a wide irregular lumen, and are rapidly reabosrbed in a process involving the apoptosis of follicular cells. A degenerative process called follicular atresia reabsorbs vitellogenic oocytes not spawned. This process can also occur, but less frequently, in oocytes in other development stages.[23]
Some fish are hermaphrodites, having testes and ovaries either at different phases in their life cycle or, like hamlets, can be simultaneously male and female.

Sensory and nervous system

Sensory and nervous system

Dorsal view of the brain of the rainbow trout.

Central nervous system
Fish typically have quite small brains relative to body size when compared with other vertebrates, typically one-fifteenth the mass of the brain from a similarly sized bird or mammal.[16] However, some fish have relatively large brains, most notably mormyrids and sharks, which have brains of about as massive relative to body weight as birds and marsupials.[17]
The brain is divided into several regions. At the front are the olfactory lobes, a pair of structure the receive and process signals from the nostrils via the two olfactory nerves.[16] The olfactory lobes are very large in fishes that hunt primarily by smell, such as hagfish, sharks, and catfish. Behind the olfactory lobes is the two-lobed telencephalon, the equivalent structure to the cerebrum in higher vertebrates. In fishes the telencephalon is concerned mostly with olfaction.[16] Together these structures form the forebrain.
Connecting the forebrain to the midbrain is the diencephalon (in the adjacent diagram, this structure is below the optic lobes and consequently not visible). The diencephalon performs a number of functions associated with hormones and homeostasis.[16] The pineal body lies just above the diencephalon. This structure performs many different functions including detecting light, maintaining circadian rhythms, and controlling colour changes.[16]
The midbrain or mesencephalon contains the two optic lobes. These are very large in species that hunt by sight, such as rainbow trout and cichlids.[16]
The hindbrain or metencephalon is particularly involved in swimming and balance.[16] The cerebellum is a single-lobed structure that is usually very large, typically the biggest part of the brain.[16] Hagfish and lampreys have relatively small cerebellums, but at the other extreme the cerebellums of mormyrids are massively developed and apparently involved in their electrical sense.[16]
The brain stem or myelencephalon is the most posterior part of the brain.[16] As well as controlling the functions of some of the muscles and body organs, in bony fish at least the brain stem is also concerned with respiration and osmoregulation.[16]

Sense organs
Most fish possess highly developed sense organs. Nearly all daylight fish have well-developed eyes that have color vision that is at least as good as a human's. Many fish also have specialized cells known as chemoreceptors that are responsible for extraordinary senses of taste and smell. Although they have ears in their heads, many fish may not hear sounds very well. However, most fishes have sensitive receptors that form the lateral line system. The lateral line system allows for many fish to detect gentle currents and vibrations, as well as to sense the motion of other nearby fish and prey.[18] Some fishes such as catfishes and sharks, have organs that detect low levels electric current.[19] Other fish, like the electric eel, can produce their own electricity.

Pain reception in fish
In 2003, Scottish scientists at the University of Edinburgh performing research on rainbow trout concluded that fish exhibit behaviors often associated with pain.[20] Professor James D. Rose of the University of Wyoming critiqued the study, claiming it was flawed.[21] Rose had published his own study a year earlier arguing that fish cannot feel pain as they lack the appropriate neocortex in the brain.[

Respiratory system

Respiratory system
Most fish exchange gases by using gills that are located on either side of the pharynx. Gills are made up of threadlike structures called filaments. Each filament contains a network of capillaries that allow a large surface area for the exchange of oxygen and carbon dioxide. Fish exchange gases by pulling oxygen-rich water through their mouths and pumping it over their gill filaments. The blood in the capillaries flows in the opposite direction to the water, causing counter current exchange. They then push the oxygen-poor water out through openings in the sides of the pharynx. Some fishes, like sharks and lampreys, possess multiple gill openings. However, most fishes have a single gill opening on each side of the body. This opening is hidden beneath a protective bony cover called an operculum.
Juvenile bichirs have external gills, a very primitive feature that they hold in common with larval amphibians.

Swim bladder of a Rudd (Scardinius erythrophthalmus)
Many fish can breathe air. The mechanisms for doing so are varied. The skin of anguillid eels may be used to absorb oxygen. The buccal cavity of the electric eel may be used to breathe air. Catfishes of the families Loricariidae, Callichthyidae, and Scoloplacidae are able to absorb air through their digestive tracts.[14] Lungfish and bichirs have paired lungs similar to those of tetrapods and must rise to the surface of the water to gulp fresh air in through the mouth and pass spent air out through the gills. Gar and bowfin have a vascularised swim bladder that is used in the same way. Loaches, trahiras, and many catfish breathe by passing air through the gut. Mudskippers breathe by absorbing oxygen across the skin (similar to what frogs do). A number of fishes have evolved so-called accessory breathing organs that are used to extract oxygen from the air. Labyrinth fish (such as gouramis and bettas) have a labyrinth organ above the gills that performs this function. A few other fish have structures more or less resembling labyrinth organs in form and function, most notably snakeheads, pikeheads, and the Clariidae family of catfish.
Being able to breathe air is primarily of use to fish that inhabit shallow, seasonally variable waters where the oxygen concentration in the water may decline at certain times of the year. At such times, fishes dependent solely on the oxygen in the water, such as perch and cichlids, will quickly suffocate, but air-breathing fish can survive for much longer, in some cases in water that is little more than wet mud. At the most extreme, some of these air-breathing fish are able to survive in damp burrows for weeks after the water has otherwise completely dried up, entering a state of aestivation until the water returns.

Tuna gills inside of the head. The fish head is oriented snout-downwards, with the view looking towards the mouth.
Fish can be divided into obligate air breathers and facultative air breathers. Obligate air breathers, such as the African lungfish, must breathe air periodically or they will suffocate. Facultative air breathers, such as the catfish Hypostomus plecostomus, will only breathe air if they need to and will otherwise rely solely on their gills for oxygen if conditions are favourable. Most air breathing fish are not obligate air breathers, as there is an energetic cost in rising to the surface and a fitness cost of being exposed to surface predators.[14]

Anatomy

Digestive system
The advent of jaws allowed fish to eat a much wider variety of food, including plants and other organisms. In fish, food is ingested through the mouth and then broken down in the esophagus. When it enters the stomach, the food is further broken down and, in many fish, further processed in fingerlike pouches called pyloric caeca. The pyloric caeca secrete digestive enzymes and absorb nutrients from the digested food. Organs such as the liver and pancreas add enzymes and various digestive chemicals as the food moves through the digestive tract. The intestine completes the process of digestion and nutrient absorptionDigestive system
The advent of jaws allowed fish to eat a much wider variety of food, including plants and other organisms. In fish, food is ingested through the mouth and then broken down in the esophagus. When it enters the stomach, the food is further broken down and, in many fish, further processed in fingerlike pouches called pyloric caeca. The pyloric caeca secrete digestive enzymes and absorb nutrients from the digested food. Organs such as the liver and pancreas add enzymes and various digestive chemicals as the food moves through the digestive tract. The intestine completes the process of digestion and nutrient absorption

Classification

Fish are a paraphyletic group: that is, any clade containing all fish also contains the tetrapods, which are not fish. For this reason, groups such as the "Class Pisces" seen in older reference works are no longer used in formal classifications.
Fish are classified into the following major groups:
Subclass Pteraspidomorphi (early jawless fish)
Class Thelodonti
Class Anaspida
(unranked) Cephalaspidomorphi (early jawless fish)
(unranked) Hyperoartia
Petromyzontidae (lampreys)
Class Galeaspida
Class Pituriaspida
Class Osteostraci
Infraphylum Gnathostomata (jawed vertebrates)
Class Placodermi (armoured fishes, extinct)
Class Chondrichthyes (cartilaginous fish)
Class Acanthodii (spiny sharks, extinct)
Superclass Osteichthyes (bony fish)
Class Actinopterygii (ray-finned fish)
Class Sarcopterygii (lobe-finned fish)
Subclass Coelacanthimorpha (coelacanths)
Subclass Dipnoi (lungfish)
Some palaeontologists consider that Conodonta are chordates, and so regard them as primitive fish. For a fuller treatment of classification, see the vertebrate article.
The various fish groups taken together account for more than half of the known vertebrates. There are almost 28,000 known extant species of fish, of which almost 27,000 are bony fish, with the remainder being about 970 sharks, rays, and chimeras and about 108 hagfishes and lampreys.[12] A third of all of these species are contained within the nine largest families; from largest to smallest, these families are Cyprinidae, Gobiidae, Cichlidae, Characidae, Loricariidae, Balitoridae, Serranidae, Labridae, and Scorpaenidae. On the other hand, about 64 families are monotypic, containing only one species. It is predicted that the eventual number of total extant species will be at least 32,500.

Definition of Fish

The term "fish" is most precisely used to describe any non-tetrapod chordate, (i.e., an animal with a backbone), that has gills throughout life and has limbs, if any, in the shape of fins.[1] Unlike groupings such as birds or mammals, fish are not a single clade but a paraphyletic collection of taxa, including hagfishes, lampreys, sharks and rays, ray-finned fishes, coelacanths, and lungfishes.[2][3]
A typical fish is ectothermic; has a streamlined body that allows it to swim rapidly; extracts oxygen from the water using gills or an accessory breathing organ to enable it to breath atmospheric oxygen; has two sets of paired fins, usually one or two (rarely three) dorsal fins, an anal fin, and a tail fin; has jaws; has skin that is usually covered with scales; and lays eggs that are fertilized internally or externally.

Fish come in many shapes and sizes. This is a sea dragon, a close relative of the seahorse. Their leaf-like appendages enable them to blend in with floating seaweed
To each of these there are exceptions. Tuna, Swordfish, and some species of sharks show some warm-blooded adaptations, and are able to raise their body temperature significantly above that of the ambient water surrounding them.[4] Streamlining and swimming performance varies from highly streamlined and rapid swimmers which are able to reach 10-20 body-lengths per second (such as tuna, salmon, and jacks) through to slow but more maneuverable species such as eels and rays that reach no more than 0.5 body-lengths per second.[5] Many groups of freshwater fish extract oxygen from the air as well as from the water using a variety of different structures. Lungfish have paired lungs similar to those of tetrapods, gouramis have a structure called the labyrinth organ that performs a similar function, while many catfish, such as Corydoras extract oxygen via the intestine or stomach.[6] Body shape and the arrangement of the fins is highly variable, covering such seemingly un-fishlike forms as seahorses, pufferfish, anglerfish, and gulpers. Similarly, the surface of the skin may be naked (as in moray eels), or covered with scales of a variety of different types usually defined as placoid (typical of sharks and rays), cosmoid (fossil lungfishes and coelacanths), ganoid (various fossil fishes but also living gars and bichirs, cycloid, and ctenoid (these last two are found on most bony fish.[7] There are even fishes that spend most of their time out of water. Mudskippers feed and interact with one another on mudflats and are only underwater when hiding in their burrows.[8] The catfish Phreatobius cisternarum lives in underground, phreatic habitats, and a relative lives in waterlogged leaf litter.[9][10]
Fish range in size from the 16 m (51 ft) whale shark to the 8 mm (just over ¼ of an inch) long stout infantfish.
Many types of aquatic animals commonly referred to as "fish" are not fish in the sense given above; examples include shellfish, cuttlefish, starfish, crayfish and jellyfish. In earlier times, even biologists did not make a distinction - sixteenth century natural historians classified also seals, whales, amphibians, crocodiles, even hippopotamuses, as well as a host of aquatic invertebrates, as fish.[11] In some contexts, especially in aquaculture, the true fish are referred to finfish (or fin fish) to distinguish them from these other animals.